Stephen (Ash) Bullard
Associate Professor



Office Location:  
324 Upchurch Hall 

Mailing Address:
Department of Fisheries
203 Swingle Hall 
Auburn University, AL 36849
Tel: (334) 844-9278
Fax: (334) 844-9208
Email Stephen (Ash) Bullard
Aquatic Parasitology Lab


Ph.D. - Gulf Coast Research Laboratory (2007)
M.Sc. - University of Southern Mississippi (2002)
B.Sc. - Cum Laude, University of South Carolina (1997)





Parasitology, diseases of fishes and aquatic invertebrates, host-parasite interactions, taxonomy & systematics, parasite & pathogen biodiversity.





Borucinska, J. & S. A. Bullard. 2011. Lesions associated with plerocerci (Platyhelminthes: Cestoda: Trypanorhyncha) in the gastric wall of a cownose ray, Rhinoptera bonasus (Myliobatiformes: Rhinopteridae) from the Northern Gulf of Mexico. Journal of Fish Diseases 34:149–157.

Bullard, S. A. 2010. Littorellicola billhawkinsi n. gen., n. sp. (Digenea: Aporocotylidae) from the myocardial lacunae of Florida pompano, Trachinotus carolinus (Carangidae) in the Gulf of Mexico; with a comment on interrelationships and functional morphology of intertrabecular aporocotylids. Parasitology International 59:587–598.

Bullard, S. A. 2010. A new species of Cardicola Short, 1953 (Digenea: Aporocotylidae) from the heart and branchial vessels of two surfperches (Perciformes: Embiotocidae) in the Eastern Pacific Ocean off California. Journal of Parasitology 96:382–388. 

Kritsky, D. C., M. Bakenhaster, E. Fajer-Avila, & S. A. Bullard. 2010. Rhabdosynochus spp. (Monogenoidea: Diplectanidae) infesting the gill lamellae of snooks, Centropomus spp. (Perciformes: Centropomidae), in Florida, and redescription of the type species, R. rhabdosynochus. Journal of Parasitology 96(5):879–886.

Jensen, K. & S. A. Bullard. 2010. Characterization of a diversity of tetraphyllidean and rhinebothriidean cestode larval types, with comments on host associations and life-cycles. International Journal for Parasitology 40:889–910.

Bullard, S. A., K. Jensen, & R. M. Overstreet. 2009. Historical account of the two family-group names in use for the single accepted family comprising the “fish blood flukes.” Acta Parasitologica 54(1):78–84.

Kritsky, D. C., E. F. Mendoza-Franco, S.A. Bullard, & V. M. Vidal-Martínez. 2009. Revision of the amphiamerican Neotetraonchus Bravo-Hollis, 1968 (Monogenoidea, Dactylogyridae), with a description of N. vegrandis n. sp. from the gill lamellae of the blue sea catfish, Ariopsis guatemalensis (Siluriformes: Ariidae), off the Pacific coast of Mexico. Systematic Parasitology 74:1–15.

Bullard, S. A. & K. Jensen. 2008. Blood flukes (Digenea: Aporocotylidae) of stingrays (Myliobatiformes: Dasyatidae): Orchispirium heterovitellatum from Himantura imbricata in the Bay of Bengal and a new genus and species from Dasyatis sabina in the Northern Gulf of Mexico. Journal of Parasitology 94:1,311–1,321.

Bullard, S. A. & R. M. Overstreet. 2008. Digeneans as enemies of fishes. In: Fish Diseases. Jorge Eiras, Helmut Segner, Thomas Wahil, and B. G. Kapoor (eds.). Science Publishers, US, pages 817–976 (invited review, editor reviewed).

Bullard, S. A., S. D. Snyder, K. Jensen, & R. M. Overstreet. 2008. New genus and species of Aporocotylidae (Digenea) from a lower actinopterygian, the American paddlefish, Polyodon spathula, (Acipenseriformes: Polyodontidae) from the Mississippi Delta. Journal of Parasitology 94:487–495.

Ulicny, K. J., D. R. Stewart, A. McElwain, E. R. Salmon, S. A. Bullard, J. O. Cook, G. B. Skomal, H. L. Pratt, Jr., & G. W. Benz. 2007. Sealing Whirl-Pak®-type Bags Containing Wet Samples. Journal of the Tennessee Academy of Sciences 82(3–4): 88–89.

Benz, G. W., B. E. Smith, S. A. Bullard, & J. S. Braswell. 2007. New genus and species of eudactylinid (Siphonstomatoida: Copepoda) from gill lamellae of ornate eagle rays, Aetomylaeus vespertilio (Myliobatidae), collected in the Beagle Gulf off Northern Australia. Journal of Parasitology 93:32–38.

Bullard, S. A., R. M. Overstreet, & J. K. Carlson. 2006. Selachohemecus benzi n. sp. (Digenea: Sanguinicolidae) from the blacktip shark Carcharhinus limbatus in the Northern Gulf of Mexico. Systematic Parasitology 63:143–154.

Bullard, S. A. & R. M. Overstreet. 2006. Psettarium anthicum sp. n. (Digenea: Sanguinicolidae) from the heart of cobia Rachycentron canadum (Rachycentridae) in the northern Gulf of Mexico. Folia Parasitologica 53:117–124 

Bullard, S. A., R. Payne, & J. S. Braswell. 2004. New genus with two new species of capsalid monogeneans from dasyatids in the Gulf of California. Journal of Parasitology 90(6):1,412–1,427.

Bullard, S. A., S. M. Dippenaar, E. R. Hoffmayer, & G. W. Benz. 2004. New locality records for Dermophthirius carcharhini MacCallum, 1926 (Monogenea: Microbothriidae) and D. maccallumi Watson and Thorson, 1976 and a list of hosts and localities for species of Dermophthirius MacCallum, 1926. Comparative Parasitology 71(1):78–80.

Bullard, S. A. & R. M. Overstreet. 2004. Two new species of Cardicola (Digenea: Sanguinicolidae) in drums (Sciaenidae) from Mississippi and Louisiana. Journal of Parasitology 90(1):128–136.

Bullard, S. A., R. Goodwin, R. Goldstein, & R. M. Overstreet. 2004. Cardicola forsteri (Digenea: Sanguinicolidae) from the heart of a northern bluefin tuna, Thunnus thynnus (Scombridae), in the Northwest Atlantic Ocean. Comparative Parasitology 71(2):245–246.

Frasca, S. Jr., V. L. Kirsipuu, S. Russell, S. A. Bullard, & G. W. Benz. 2004. Opercular lesions in black drum (Pogonias cromis) associated with infections of the sea louse Sciaenophilus tenuis (Copepoda: Caligidae). Acta Ichthyologica et Piscatoria 34(2):115–127.

Blaylock, R. B., S. A. Bullard, & C. M. Whipps. 2004. Kudoa hypoepicardialis sp. n. (Myxozoa: Kudoidae) and associated lesions from the heart of seven perciform fishes in the northern Gulf of Mexico. Journal of Parasitology 90(3):584–593.

Benz, G. W., R. Hocking, A. Kowunna, Sr., S. A. Bullard, J. C. George. 2004. A second species of Arctic shark: Pacific sleeper shark Somniosus pacificus from Point Hope Alaska. Polar Biology 27:250–252.

Benz, G. W. & S. A. Bullard. 2004. Metazoan parasites and associates of chondrichthyans with emphasis on taxa harmful to captive hosts. In: The husbandry of elasmobranch fishes. Smith, M. F. L., D. I. Warmolts, D. A. Thoney, and R. E. Hueter (eds.). Special Publication No. 16, Ohio Biological Survey, Columbus, OH. Pages 325–416.

Bullard, S. A. & S. M. Dippenaar. 2003. Branchotenthes robinoverstreeti gen. and sp. n. (Monogenea: Hexabothriidae) from gill filaments of the bowmouth guitarfish, Rhina ancylostoma (Rhynchobatidae), in the Indian Ocean. Journal of Parasitology 89(3):595–601.

Bullard, S. A., R. Hocking, R. J. Goldstein, & J. Jewell. 2003. A new locality record and three new host records for Neobenedenia melleni (Monogenea: Capsalidae). Gulf and Caribbean Research 15:1–4. 

Bullard, S. A. & R. M. Overstreet. 2003. Elaphrobates euzeti gen. and sp. n. (Digenea: Sanguinicolidae) from snappers (Lutjanidae) in the Gulf of Mexico. In: Taxonomie, écologie et évolution des métazoaires parasites. Taxonomy, ecology and evolution of metazoan parasites. (Livre hommage à Louis Euzet). Tome 1. Combes C. and Jourdane J. (eds), PUP Perpignan:97–113 (peer-reviewed).

Benz, G. W., S. A. Bullard, & A. D. M. Dove. 2002. Metazoan parasites of fishes: Synoptic information for aquarists. Proceedings of the Meeting of the American Zoologists and Aquarists, Chattanooga, TN (non-peer reviewed).

Bullard, S. A. & R. M. Overstreet. 2002. Potential pathological effects of blood flukes (Digenea: Sanguinicolidae) on pen-reared marine fishes. Proceedings of the 53rdGulf and Caribbean Fisheries Institute: pages 10–25 (editor reviewed).

Benz, G. W., B. E. Smith, & S. A. Bullard. 2001. Kroyerina deetsorum n. sp. (Copepoda: Kroyeriidae) from the olfactory sacs of Atlantic sharpnose sharks (Rhizoprionodon terraenovae) captured in the Gulf of Mexico and Northwestern Atlantic and first report of copepodids representing Kroyeriidae. Journal of Parasitology 87(6):1279–1290.

Bullard, S. A., S. Frasca Jr., & G. W. Benz. 2001. Gill lesions associated with Erpocotyle tiburonis (Monogenea: Hexabothriidae) on wild and aquarium-held bonnethead sharks (Sphyrna tiburo). Journal of Parasitology 87(5):972–977.

Benz, W. W., S. A. Bullard, & A. D. M. Dove. 2001. Metazoan parasites of fishes: synoptic information and portal to the literature for aquarists.  Pp. 1-15.  In: Regional Conference Proceedings 2001. Anonymous (ed.). American Zoo and Aquarium Association, Silver Spring, MD. (non-peer reviewed proceedings)

Bullard, S. A., S. Frasca, Jr., & G. W. Benz. 2000. Skin lesions caused by Dermophthirius penneri (Monogenea: Microbothriidae) on wild-caught blacktip sharks (Carcharhinus limbatus). Journal of Parasitology 86(3):618–622. 

Bullard, S. A., G. W. Benz, R. M. Overstreet, E. H. Williams, Jr., & J. Hemdal. 2000. Six new host records and updated list of wild hosts for Neobenedenia melleni (McCallum, 1927) (Monogenea: Capsalidae). Comparative Parasitology 67(2):190–196.

Bullard, S. A. & R. M. Overstreet. 2000. Calicotyle californiensis n. sp. and Calicotyle urobati n. sp. (Monogenea: Calicotylinae) from elasmobranchs in the Gulf of California. Journal of Parasitology 86(5):939–944.

Bullard, S. A., G. W. Benz, & J. S. Braswell. 2000. Dionchus postoncomiracidia (Monogenea: Dionchidae) from skin of blacktip sharks, Carcharhinus limbatus (Carcharhinidae). Journal of Parasitology 86(2):245–250.

Benz, G. W., Z. Kabata, & S. A. Bullard. 2000. Margolisius abditus n. gen., n. sp. (Copepoda: Lernaeopodidae) from gill lamellae of a remora (Remora remora) collected in the Gulf of California. Journal of Parasitology 86(2):241–244.






Undergraduate Graduate
Fish Anatomy and Physiology - FISH5440 Fish Anatomy and Physiology - FISH6440
Marine Ichthyology - FISH5725 Marine Ichthyology - FISH6725
Special Problems in Parasitology - FISH4970 Special Problems in Parasitology - FISH7970
Special Problems in Pathology of Aquatic Organisms - FISH7960


Last updated: 04/25/2014